Arizona Ants
Ants are a critical component of the earth’s terrestrial ecosystems. They consume and break down large amounts of material, they control the populations of numerous species, and they provide food for many others. For instance, ants make up 40% of the diet of the Northern Flicker, a common Arizona bird. Despite being small and not so visible, ants account for 15% to 25% of all animal biomass on our planet’s land surface—far more than any other animal group.
So how much do the ants weigh? Using guesstimates from the references I made assumptions that the average ant weighs .004 grams, and that there are 1,000,000 ants per human. Thus, the earth’s ants weigh around 21 billion tons (I didn’t try to use dry weight).
Not to be beaten by little bugs, however, humanity is increasing its weight at a rapid rate. Issac Asimov once estimated that if our numbers continued to grow at the 1970’s rate of 2% (doubling in 35 years), the mass of humanity would match that of the whole planet in about 1500 years (Asimov 1975). Probably no ants (or any other animal) could survive that. Of course, our birth rate is declining, and even though our life expectancy is increasing, doubling our numbers will take longer and longer. If we assume that the average person weighs 120 pounds, our species currently weighs 420 million tons. If we double every 100 years, and if we do not destroy any ant habitat (OK, that is not a reasonable assumption. Nevertheless…), it will take 600 years for us to become heavier than the ants. After that, we get ahead and stay ahead.
The photo shows Myrmecocystus navajo, a member of the Formicinae subfamily. This one was collected in Yavapai County, Arizona. The photo may be by Barry Bolton, but it could be by Stefan Cover or Bob Johnson. It is from the AntWeb website.
_______
Arizona has 318 ant species, more than any other U. S. state. Ants are found from the lowest desert areas to near the tops of the highest mountains. Harvester Ants (Pogonomyrmex, Messor, and Pheidole) are most abundant in warm desert areas. Carpenter (Camponotus) and Wood Ants (Formica) are more common in cooler uplands and mountains.
Some ants are quite secretive, and either hide or camouflage their nests. Many ants have a painful bite or sting, but some use other means of defense. Once while studying ants near Red Pine Mountain, my friend Ray Turner was using an aspirator to capture ants from a camouflaged Formica nest when he suddenly began gasping and momentarily could not catch his breath. Ray was nearly overcome by the formic acid odor released by the frightened ants. Some birds place these ants in their feathers to repel parasites.
According to Stefan Cover and Bob Johnson (www.antweb.org/arizona.jsp), 12 of Arizona’s ant species are introduced, but none of these are considered to be problems. Fire Ants from South America (Solenopsis invicta) have been found in Arizona, but the only known colony has been eradicated. Fire Ants are highly destructive. They form large colonies that displace other ants, alter habitats, and consume native animals and their offspring (Arizona-Sonora Desert Museum). Edward O. Wilson once stuck his hand in a Fire Ant nest to observe the response. He was stung more than 1,000 times.
Fire ant stings are reported to be quite painful, but according to Justin O. Schmidt (1990), they are only “mildly alarming” in comparison to some others. For instance, he describes the Bullet ant sting as “Pure, intense, brilliant pain. Like fire-walking over flaming charcoal with a 3-inch rusty nail in your heel” (quoted in Wikipedia in the article on the “Schmidt Sting Pain Index”).
The Ants (Holldobler and Wilson 1990), Ant Ecology (see Alonso 2010) and the novel, Ant Hill (Wilson 2010) provide extensive information about ants. Don’t miss the article by Berenbaum (2003). It gives an in-depth discussion of the pain caused by ant stings, and it is a delightful example of creative nonfiction. Antweb provides photographs and field guides for Arizona ants.
Ants, like some wise people, eschew cold weather. While they are out and about next summer we can discuss collecting, identifying, and photographing them.
Conservation status
There would be enormous costs to our planetary ecosystems if ant habitat continues to be lost to human development. Some scientists have convincingly argued that ants should be at the center of conservation concerns. Perhaps because of their vast numbers and ubiquity, ants have, in fact, been given relatively little attention. In a recent article on ant ecology, Leeanne Alonso (2009) discusses the approaches that need to be taken for effective ant conservation. As Wilson points out in the forward, however, “We have only begun to explore the full impact of ants on the natural ecosystems of the planet…” (viii).
| Numbers (species identified/probable total) | |||
| World | US | AZ | T&E |
| 12500/22000* | 700(?) | 318/334** | |
| *Wikipedia. **species/subspecies, Antweb. | |||
References
Allred, D.M. 1982. Ants of Utah. Great Basin Naturalist 42: 415-511.
Alonso, L.E. 2010. Ant conservation: Current status and a call to action. Chapter 4 in L. Lach, C.L. Parr, and K.L. Abbott. Ant Ecology. Forward by E.O. Wilson. Oxford University Press, New York, NY. 409 p.
Asimov, I. 1975. Science past–science future. Doubleday, Garden City, New York, NY. 346 p.
Antweb: www.antweb.org/arizona.jsp. Principal authors: Stefan Cover and Bob Johnson.
Arizona-Sonora Desert Museum: http://www.desertmuseum.org/invaders/invaders_fireant.php.
Berenbaum, M. 2003. A Stinging Commentary. American Entomologist 49: 68-69.
Holldobler, B., and E.O. Wilson. 1990. The ants. Belknap Press, New York, NY. 746 p.
Johnson, R.A. 1996. Arizona ants. Arizona Wildlife Views, June 1996: 3-5.
Schmidt, J. O. 1990. Hymenoptera venoms: striving toward the ultimate defense against vertebrates, pp. 387–419 in D.L. Evans and J.O. Schmidt, eds., Insect defenses: adaptive mechanisms and strategies of prey and predators. State University of New York Press, Albany, NY.
Wilson, E. O. 2010. Ant Hill. Norton, New York, NY. 382 p.
Checklist
The field guides provided by AntWeb can be used as checklists, but the one below is more compact because it is merely a list—no help with identification. It does sort species into subfamilies. Many species occur in the rich southeastern Arizona desert and uplands far from the Agua Fria River Basin. Probably half the species in the list below do not occur in the Basin, but I could not find any lists for any region smaller than the state.
|
Arizona Ants |
|
| Amblyoponinae | |
| Amblyopone orizabana | |
| Amblyopone pallipes | |
| Cerapachyinae | |
| Acanthostichus arizonensis | |
| Acanthostichus punctiscapus | |
| Cerapachys augustae | |
| Dolichoderinae | |
| Dorymyrmex az01 (Cone or odorous ant)) | |
| Dorymyrmex az02 | |
| Dorymyrmex az03 | |
| Dorymyrmex az04 | |
| Dorymyrmex az05 | |
| Dorymyrmex az06 | |
| Dorymyrmex bicolor | |
| Dorymyrmex flavus | |
| Dorymyrmex insanus | |
| Dorymyrmex wheeleri | |
| Forelius az01 | |
| Forelius az02 | |
| Forelius mccooki | |
| Linepithema humile | |
| Liometopum apiculatum | |
| Liometopum luctuosum (Pine tree ant) | |
| Tapinoma sessile | |
| Ecitoninae (Army Ants) | |
| Neivamyrmex agilis | |
| Neivamyrmex andrei | |
| Neivamyrmex az01 | |
| Neivamyrmex az02 | |
| Neivamyrmex az03 | |
| Neivamyrmex fuscipennis | |
| Neivamyrmex graciellae | |
| Neivamyrmex harrisii | |
| Neivamyrmex kiowapache | |
| Neivamyrmex leonardi | |
| Neivamyrmex melshaemeri | |
| Neivamyrmex microps | |
| Neivamyrmex minor | |
| Neivamyrmex nigrescens | |
| Neivamyrmex nyensis cf | |
| Neivamyrmex opacithorax | |
| Neivamyrmex rugulosus | |
| Neivamyrmex swainsonii | |
| Neivamyrmex texanus | |
| Ectatomminae | |
| Typhlomyrmex az01 | |
| Formicinae (Carpenter Ants) | |
| Acropyga epedana | |
| Brachymyrmex depilis | |
| Brachymyrmex obscurior | |
| Camponotus festinatus | |
| Camponotus fragilis | |
| Camponotus hyatti | |
| Camponotus laevigatus | |
| Camponotus microps | |
| Camponotus mina | |
| Camponotus modoc | |
| Camponotus ocreatus | |
| Camponotus papago | |
| Camponotus pudorosus | |
| Camponotus sansabeanus | |
| Camponotus sayi | |
| Camponotus schaefferi | |
| Camponotus semitestaceus | |
| Camponotus trepidulus | |
| Camponotus ulcerosus | |
| Camponotus vafer | |
| Camponotus vicinus | |
| Formica alpina | |
| Formica altipetens | |
| Formica argentea | |
| Formica aserva | |
| Formica az01 | |
| Formica az02 | |
| Formica az03 | |
| Formica az04 | |
| Formica canadensis | |
| Formica ciliata | |
| Formica comata | |
| Formica dakotensis | |
| Formica densiventris | |
| Formica foreliana | |
| Formica fusca marcida | |
| Formica gnava | |
| Formica gynocrates | |
| Formica lasioides | |
| Formica moki | |
| Formica neoclara | |
| Formica neogagates | |
| Formica neorufibarbis | |
| Formica obscuripes | |
| Formica obtusopilosa | |
| Formica occidua | |
| Formica occulta | |
| Formica opaciventris | |
| Formica oreas | |
| Formica perpilosa | |
| Formica podzolica | |
| Formica propinqua | |
| Formica puberula | |
| Formica subnitens | |
| Formica wheeleri | |
| Formica xerophila | |
| Lasius alienus | |
| Lasius arizonicus | |
| Lasius az01 | |
| Lasius colei | |
| Lasius coloradensis | |
| Lasius crypticus | |
| Lasius fallax | |
| Lasius flavus | |
| Lasius humilis | |
| Lasius interjectus | |
| Lasius latipes | |
| Lasius murphyi | |
| Lasius niger | |
| Lasius occidentalis | |
| Lasius pallitarsis | |
| Lasius sitiens | |
| Lasius subumbratus | |
| Lasius umbratus | |
| Myrmecocystus (Honey Ants) az01 | |
| Myrmecocystus az02 | |
| Myrmecocystus depilis | |
| Myrmecocystus flaviceps | |
| Myrmecocystus kennedyi | |
| Myrmecocystus mendax | |
| Myrmecocystus mexicanus | |
| Myrmecocystus mimicus | |
| Myrmecocystus navajo | |
| Myrmecocystus placodops | |
| Myrmecocystus romainei | |
| Myrmecocystus tenuinodis | |
| Myrmecocystus yuma | |
| Nylanderia austroccidua | |
| Nylanderia bruesii | |
| Nylanderia terricola | |
| Nylanderia vividula | |
| Paratrechina longicornis | |
| Polyergus breviceps | |
| Prenolepis imparis | |
| Myrmicinae (Harvester Ants) | |
| Acromyrmex versicolor | |
| Aphaenogaster albisetosa | |
| Aphaenogaster boulderensis | |
| Aphaenogaster cockerelli | |
| Aphaenogaster huachucana | |
| Aphaenogaster megommata | |
| Aphaenogaster occidentalis | |
| Aphaenogaster punctaticeps | |
| Atta mexicana | |
| Cardiocondyla mauritanica | |
| Cardiocondyla minutior | |
| Cardiocondyla venustula | |
| Cephalotes rohweri | |
| Crematogaster az01 | |
| Crematogaster az02 | |
| Crematogaster browni | |
| Crematogaster colei | |
| Crematogaster dentinodis | |
| Crematogaster depilis | |
| Crematogaster emeryana | |
| Crematogaster hespera | |
| Crematogaster isolata | |
| Crematogaster larreae | |
| Crematogaster navajoa | |
| Crematogaster nocturna | |
| Crematogaster opaca | |
| Crematogaster opuntiae | |
| Crematogaster smithi | |
| Crematogaster torosa | |
| Cyphomyrmex flavidus | |
| Cyphomyrmex wheeleri | |
| Dolopomyrmex pilatus | |
| Leptothorax az01 | |
| Leptothorax az02 | |
| Leptothorax crassipilis | |
| Messor az01 (Seed harvesters) | |
| Messor lobognathus | |
| Messor pergandei | |
| Messor smithi | |
| Monomorium pharaonis | |
| Monomorium az01 | |
| Monomorium az02 | |
| Monomorium az03 | |
| Monomorium emersoni | |
| Myrmecina americana | |
| Myrmecina cryptica | |
| Myrmica az01 | |
| Myrmica az02 | |
| Myrmica az03 | |
| Myrmica az04 | |
| Myrmica az05 | |
| Myrmica az06 | |
| Myrmica az07 | |
| Myrmica az08 | |
| Myrmica discontinua | |
| Myrmica incompleta | |
| Myrmica magniceps | |
| Myrmica rugiventris | |
| Myrmica striolagaster | |
| Myrmica tahoensis | |
| Myrmica wheeleri | |
| Pheidole artemisia | |
| Pheidole az01 | |
| Pheidole az02 | |
| Pheidole az03 | |
| Pheidole az04 | |
| Pheidole az05 | |
| Pheidole az06 | |
| Pheidole az07 | |
| Pheidole az08 | |
| Pheidole az09 | |
| Pheidole az10 | |
| Pheidole az11 | |
| Pheidole az12 | |
| Pheidole az13 | |
| Pheidole az14 | |
| Pheidole az15 | |
| Pheidole barbata | |
| Pheidole bicarinata | |
| Pheidole californica | |
| Pheidole cavigenis | |
| Pheidole cerebrosior | |
| Pheidole ceres | |
| Pheidole clydei | |
| Pheidole cockerelli | |
| Pheidole coloradensis | |
| Pheidole desertorum | |
| Pheidole diversipilosa | |
| Pheidole floridana | |
| Pheidole furtiva | |
| Pheidole gilvescens | |
| Pheidole hyatti | |
| Pheidole juniperae | |
| Pheidole marcidula | |
| Pheidole micula | |
| Pheidole militicida | |
| Pheidole obtusospinosa | |
| Pheidole paiute | |
| Pheidole perpilosa | |
| Pheidole portalensis | |
| Pheidole psammophila | |
| Pheidole rhea | |
| Pheidole rugulosa | |
| Pheidole sciara | |
| Pheidole sciophila | |
| Pheidole senex | |
| Pheidole soritis | |
| Pheidole spadonia | |
| Pheidole tepicana | |
| Pheidole tetra | |
| Pheidole titanis | |
| Pheidole tysoni | |
| Pheidole vallicola | |
| Pheidole virago | |
| Pheidole vistana | |
| Pheidole xerophila | |
| Pogonomyrmex anergismus (Seed harvesters) | |
| Pogonomyrmex apache | |
| Pogonomyrmex barbatus | |
| Pogonomyrmex bicolor | |
| Pogonomyrmex californicus | |
| Pogonomyrmex colei | |
| Pogonomyrmex desertorum | |
| Pogonomyrmex huachucanus | |
| Pogonomyrmex imberbiculus | |
| Pogonomyrmex magnacanthus | |
| Pogonomyrmex maricopa | |
| Pogonomyrmex occidentalis | |
| Pogonomyrmex pima | |
| Pogonomyrmex rugosus | |
| Pyramica arizonica | |
| Pyramica chiricahua | |
| Pyramica membranifera | |
| Rogeria foreli | |
| Solenopsis amblychila | |
| Solenopsis aurea | |
| Solenopsis az01 | |
| Solenopsis az02 | |
| Solenopsis az03 | |
| Solenopsis az04 | |
| Solenopsis az05 | |
| Solenopsis az06 | |
| Solenopsis invicta | |
| Solenopsis krockowi | |
| Solenopsis xyloni | |
| Stenamma californicum | |
| Stenamma chiricahua | |
| Stenamma huachucanum | |
| Stenamma snellingi | |
| Strumigenys louisianae | |
| Temnothorax andrei | |
| Temnothorax az01 | |
| Temnothorax az03 | |
| Temnothorax az04 | |
| Temnothorax az05 | |
| Temnothorax az06 | |
| Temnothorax az07 | |
| Temnothorax az08 | |
| Temnothorax ca09 | |
| Temnothorax carinatus | |
| Temnothorax emmae | |
| Temnothorax josephi | |
| Temnothorax neomexicanus | |
| Temnothorax nitens | |
| Temnothorax obliquicanthus | |
| Temnothorax obturator | |
| Temnothorax pergandei | |
| Temnothorax polita | |
| Temnothorax rugatulus | |
| Temnothorax silvestrii | |
| Temnothorax stenotyle | |
| Temnothorax tricarinatus | |
| Tetramorium hispidum | |
| Tetramorium spinosum | |
| Trachymyrmex arizonensis | |
| Trachymyrmex carinatus | |
| Trachymyrmex desertorum | |
| Trachymyrmex nogalensis | |
| Trachymyrmex pomonae | |
| Ponerinae | |
| Hypoponera az01 | |
| Hypoponera bca01 | |
| Hypoponera inexorata | |
| Hypoponera opaciceps | |
| Hypoponera opacior | |
| Hypoponera punctatissima | |
| Odontomachus clarus | |
| Pseudomyrmecinae | |
| Pseudomyrmex apache | |
|
Pseudomyrmex pallidus |
|
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